Protein Information

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UniProt AC / UniProt ID P10636 / TAU_HUMAN
Protein Name Microtubule-associated protein tau
Gene Name Name: MAPT
Synonyms:MAPTL,MTBT1,TAU;
Organism Homo sapiens (Human)

PRO ID PR:P10636
DAG View
PRO Name microtubule-associated protein tau (human)
Definition A microtubule-associated protein tau that is encoded in the genome of human.
Short Label hMAPT
Category organism-gene

Interactive Sequence View

Select/align proteoforms across species


P10636 (MAPT) as Substrate



Site PTM Type PTM Enzyme Score Source PMID
A2 Acetylation UniProt 1512244
Y18 Phosphorylation P06241 (FYN) , P43405 (SYK) neXtProt   PhosphoSitePlus   RLIMS-P+   RLIMS-P   Signor   UniProt 33363455, 28919467, 29782321, ...18070606, 20110609, 25963821, 32929078, 15356202, 27520374, 27113998, 31698056, 14999081, 29958544, 16014719, 21794954
T39 Phosphorylation PhosphoSitePlus
S46 Phosphorylation PhosphoSitePlus   RLIMS-P 33597014
T50 Phosphorylation PhosphoSitePlus
S56 Phosphorylation PhosphoSitePlus
S61 Phosphorylation PhosphoSitePlus
T123 O-Glycosylation GlyGen 20706749
S171 Phosphorylation PhosphoSitePlus
T173 Phosphorylation PhosphoSitePlus
E199 Phosphorylation IntAct 17078951
G205 Phosphorylation IntAct 17078951
P212 Phosphorylation IntAct 17078951
S214 Phosphorylation O00141 (SGK1) neXtProt   PhosphoSitePlus   RLIMS-P   Signor   UniProt 29044200, 14636947, 12387894, ...21387172, 26095350, 16982696, 20679343, 28646348, 29382817, 33192469, 26223541, 27708431, 35064472, 36203870, 9614189, 16645045, 23983461, 22003387, 24367683, 27193083, 33469985
D231 Phosphorylation IntAct 17078951
S232 Phosphorylation PhosphoSitePlus
S235 Phosphorylation Q00535 (CDK5) RLIMS-P   Signor 21215781, 22723997, 30318466, ...27641626, 24559475
S238 O-Glycosylation GlyGen 21327254
S238 Phosphorylation PhosphoSitePlus   RLIMS-P 25524708
S255 Phosphorylation Q13131 (PRKAA1) neXtProt 23583109, 21204788
S355 Phosphorylation Q13131 (PRKAA1) neXtProt   PhosphoSitePlus 23583109, 21204788
S369 Phosphorylation Q13627 (DYRK1A) neXtProt   PhosphoSitePlus 17906291, 21573099, 11311121
T386 Phosphorylation PhosphoSitePlus
S388 Phosphorylation PhosphoSitePlus   RLIMS-P 33629708
S396 Phosphorylation P49841 (GSK3B) , Q13131 (PRKAA1) , P48730 (CSNK1D) IntAct   neXtProt   RLIMS-P+   RLIMS-P   Signor   UniProt 33221742, 27609071, 10891598, ...30397132, 27406053, 23583109, 35339839, 23634895, 20831597, 25096780, 10794842, 1735956, 16519507, 30389461, 25651436, 30057671, 21204788, 36453396, 28328017, 26095350, 29889065, 20679343, 32155989, 1899488, 29844403, 25974414, 32967251, 29062069, 26502977, 28035925, 27228974, 33629708, 31479419, 26828481, 35378696, 33256579, 29958544, 16877359, 28225806, 27708431, 29914030, 34995924, 26629858, 23977276, 33328879, 14761950, 25735452, 30958368, 27221467, 22606286, 17078951, 29191219, 31824301, 31267651, 1336152, 36463652, 29670132, 24033439, 8835879, 8804716, 28634382, 34220449, 25641096, 25165879, 32815315, 32093751, 27117003, 30870531, 33218072, 27922637, 8892109, 36030903, 35698879, 35600626, 21573099, 31787873
S400 O-Glycosylation GlyGen 21327254, 20706749, 34220449
S400 Phosphorylation P49841 (GSK3B) RLIMS-P+   RLIMS-P   Signor 20679343, 27071766, 34220449, ...16519507
L404 Phosphorylation IntAct 17078951
S411 Phosphorylation PhosphoSitePlus
T449 Phosphorylation PhosphoSitePlus
S451 Phosphorylation PhosphoSitePlus
T466 Phosphorylation PhosphoSitePlus
T470 Phosphorylation PhosphoSitePlus   UniProt 9614189
T492 Phosphorylation PhosphoSitePlus
T498 Phosphorylation P53779 (MAPK10) , P45983 (MAPK8) , P49841 (GSK3B) , Q00535 (CDK5) , P45984 (MAPK9) neXtProt   PhosphoSitePlus   Signor   UniProt 15546861, 12387894, 15228592
S501 Phosphorylation PhosphoSitePlus
S502 Phosphorylation PhosphoSitePlus
S508 Phosphorylation PhosphoSitePlus
S512 Phosphorylation P49841 (GSK3B) PhosphoSitePlus   Signor 9771888
Y514 Phosphorylation PhosphoSitePlus   UniProt 16923168
S515 Phosphorylation P49841 (GSK3B) , Q5TCY1 (TTBK1) PhosphoSitePlus   Signor   UniProt 16923168, 9832145
S516 Phosphorylation Q5TCY1 (TTBK1) , Q00535 (CDK5) , P45984 (MAPK9) , P45983 (MAPK8) , P49841 (GSK3B) , P53779 (MAPK10) neXtProt   PhosphoSitePlus   Signor   UniProt 16923168, 19451179, 12387894, ...9614189, 15546861, 15228592, 20308788
S519 Phosphorylation Q13627 (DYRK1A) , P53779 (MAPK10) , Q00535 (CDK5) , P45983 (MAPK8) , P48730 (CSNK1D) , Q5TCY1 (TTBK1) , P45984 (MAPK9) , P28482 (MAPK1) , P49841 (GSK3B) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 10737616, 23186163, 15228592, ...18220336, 14761950, 16923168, 12226093, 21327254, 19451179, 12387894, 9614189, 15546861, 24275569, 17906291, 17078951, 11311121, 21573099, 21215781
T522 Phosphorylation P45983 (MAPK8) , P48730 (CSNK1D) , P49841 (GSK3B) , Q00535 (CDK5) , P28482 (MAPK1) , P53779 (MAPK10) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 12226093, 14761950, 10737616, ...19451179, 12387894, 15546861, 15228592, 20308788
S525 O-Glycosylation GlyGen   UniProt 21327254
S525 Phosphorylation PhosphoSitePlus
S527 Phosphorylation PhosphoSitePlus
T529 Phosphorylation P45984 (MAPK9) , Q8IWQ3 (BRSK2) , P45983 (MAPK8) , P49841 (GSK3B) , Q8TDC3 (BRSK1) , Q13627 (DYRK1A) , P53779 (MAPK10) , Q92630 (DYRK2) , P57059 (SIK1) , Q00535 (CDK5) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 15705853, 11181841, 17906291, ...2387894, 19451179, 12387894, 9614189, 15546861, 18599021, 15228592, 17482548, 21985311, 11311121, 21573099, 21215781
S531 Phosphorylation P49841 (GSK3B) , P57059 (SIK1) , P17612 (PRKACA) , Q00535 (CDK5) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 19451179, 12387894, 9614189, ...15546861, 9832145, 16443603, 12435421, 21985311
T534 Phosphorylation P53779 (MAPK10) , Q00535 (CDK5) , P49841 (GSK3B) , P45983 (MAPK8) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 11181841, 2387894, 19451179, ...12387894, 15228592, 17078951, 16443603
T537 Phosphorylation PhosphoSitePlus
T548 Phosphorylation Q9UQM7 (CAMK2A) , P45984 (MAPK9) , P17612 (PRKACA) , P49841 (GSK3B) , Q00535 (CDK5) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 15546861, 17287340, 10090741, ...12387894, 9565682, 15228592, 23186163, 16443603, 9735171, 14690523, 21215781
S552 Phosphorylation Q9UQM7 (CAMK2A) , Q00535 (CDK5) phospho.ELM   PhosphoSitePlus   Signor   UniProt 9565682, 17287340, 10090741, ...9614189, 12387894, 15546861, 23186163, 16443603
S554 Phosphorylation PhosphoSitePlus   UniProt 16443603, 8999860
S555 O-Glycosylation GlyGen   UniProt 21327254
S555 Phosphorylation IEDB   PhosphoSitePlus 30318466
S558 Phosphorylation PhosphoSitePlus
K571 Ubiquitination UniProt 16443603
S575 Phosphorylation phospho.ELM 12714590
S579 Phosphorylation P57059 (SIK1) , Q9P0L2 (MARK1) , P17612 (PRKACA) , Q8TDC3 (BRSK1) , P49841 (GSK3B) , P49840 (GSK3A) , Q9UQM7 (CAMK2A) , O96017 (CHEK2) , Q8IWQ3 (BRSK2) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 15705853, 8999860, 9771888, ...10090741, 19451179, 9614189, 15546861, 7706316, 17482548, 16443603, 21985311, 12714590, 23666762, 21215781, 12435421
T580 Phosphorylation PhosphoSitePlus
S602 Phosphorylation PhosphoSitePlus   UniProt 8999860
S610 Phosphorylation P57059 (SIK1) neXtProt   PhosphoSitePlus   UniProt 21985311, 7706316
K611 Methylation PhosphoSitePlus
S622 Phosphorylation P57059 (SIK1) neXtProt   PhosphoSitePlus   UniProt 21985311, 8999860, 7706316
Y627 Phosphorylation PhosphoSitePlus
K628 Acetylation PhosphoSitePlus
K628 Ubiquitination UniProt 16443603
S633 Phosphorylation PhosphoSitePlus
T636 Phosphorylation PhosphoSitePlus
S637 Phosphorylation Q16512 (PKN1) Signor 11104762
S641 Phosphorylation P49840 (GSK3A) , Q9P0L2 (MARK1) , P57059 (SIK1) neXtProt   PhosphoSitePlus   Signor   UniProt 21985311, 7706316, 21215781
S669 Phosphorylation UniProt 8999860
K670 Ubiquitination UniProt 16443603
S673 Phosphorylation Q9P0L2 (MARK1) , P17612 (PRKACA) , P57059 (SIK1) neXtProt   PhosphoSitePlus   Signor   UniProt 12435421, 21985311, 7706316, ...21215781
K686 Acetylation PhosphoSitePlus
K687 Acetylation PhosphoSitePlus
T690 Phosphorylation phospho.ELM 12387894
T694 Phosphorylation PhosphoSitePlus
Y711 Phosphorylation PhosphoSitePlus
S713 Phosphorylation P49840 (GSK3A) , Q00535 (CDK5) , P45984 (MAPK9) , P53779 (MAPK10) , P45983 (MAPK8) , P49841 (GSK3B) IEDB   neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 15546861, 23186163, 11181841, ...14761950, 21327254, 19451179, 12387894, 9614189, 9565682, 17229088, 15228592, 29670132, 16443603, 20080974, 19690332
S717 O-Glycosylation GlyGen   UniProt 20706749, 21327254
S717 Phosphorylation P49841 (GSK3B) PhosphoSitePlus   Signor   UniProt 19690332, 12387894
T720 Phosphorylation PhosphoSitePlus
S721 Phosphorylation P49841 (GSK3B) , Q00535 (CDK5) , P49840 (GSK3A) , P45983 (MAPK8) , P45984 (MAPK9) , Q13627 (DYRK1A) , P28482 (MAPK1) , P53779 (MAPK10) , P48730 (CSNK1D) neXtProt   phospho.ELM   PhosphoSitePlus   Signor   UniProt 15546861, 23186163, 11181841, ...14761950, 10737616, 21327254, 19451179, 12387894, 9614189, 9565682, 17229088, 15228592, 17906291, 21215781, 20679343, 20080974, 11311121, 21573099, 19690332
S726 Phosphorylation P17612 (PRKACA) PhosphoSitePlus   UniProt 15546861, 12435421, 19690332
S729 Phosphorylation PhosphoSitePlus
S730 Phosphorylation PhosphoSitePlus
T731 Phosphorylation PhosphoSitePlus
S733 Phosphorylation PhosphoSitePlus   UniProt 16923168
S739 Phosphorylation P53779 (MAPK10) , P45984 (MAPK9) , P45983 (MAPK8) , Q5TCY1 (TTBK1) IEDB   neXtProt   PhosphoSitePlus   Signor   UniProt 16923168, 19451179, 9614189, ...15228592, 16443603, 30001207
T744 Phosphorylation PhosphoSitePlus   UniProt 16923168
Site PTM Type PTM Enzyme Score Source PMID
S55 Phosphorylation PhosphoSitePlus
T65 Phosphorylation HPRD 12387894
S73 Phosphorylation HPRD   PhosphoSitePlus 17287340
T77 Phosphorylation HPRD   PhosphoSitePlus 17287340
T95 Phosphorylation P06493 (CDK1) HPRD 9614189
T117 Phosphorylation P28482 (MAPK1) HPRD 9832145, 10737616
T123 Phosphorylation P28482 (MAPK1) , P53778 (MAPK12) , P49841 (GSK3B) HPRD 11181841, 7566346, 9771888, ...9832145, 18510355, 9199504
S126 Phosphorylation P49841 (GSK3B) HPRD 12387894, 9771888
S137 Phosphorylation P17612 (PRKACA) , P49841 (GSK3B) HPRD 11181841, 12387894, 9771888
S140 Phosphorylation P49841 (GSK3B) , P17612 (PRKACA) HPRD 7566346, 9832145, 12387894, ...9771888
S141 Phosphorylation P17612 (PRKACA) , P49841 (GSK3B) HPRD   PRO 19413330, 9771888, 7566346, ...12387894, 9565682, 9485060
S144 Phosphorylation Q00535 (CDK5) , P49841 (GSK3B) , P28482 (MAPK1) , P53778 (MAPK12) HPRD   PRO 19413330, 9771888, 7566346, ...12226093, 1330687, 12387894, 19415658, 9565682, 9832145, 9485060, 9199504
T147 Phosphorylation P53778 (MAPK12) , P49841 (GSK3B) , Q00535 (CDK5) HPRD   PRO 19413330, 11181841, 19562805, ...9199504, 12226093, 9565682, 9485060, 9771888
S150 Phosphorylation P49841 (GSK3B) HPRD 19562805, 11311121
S152 Phosphorylation HPRD 9832145
T154 Phosphorylation P17612 (PRKACA) , P51812 (RPS6KA3) , Q00535 (CDK5) , P49841 (GSK3B) HPRD 9546672, 11181841, 12387894, ...11311121, 9832145, 9353289
S156 Phosphorylation Q16512 (PKN1) , Q00535 (CDK5) , P17612 (PRKACA) HPRD   PhosphoSitePlus 9546672, 11104762, 12387894, ...9832145, 8163474, 12435421
T159 Phosphorylation Q00535 (CDK5) , P49841 (GSK3B) HPRD 7566346, 9832145, 12387894, ...11181841
T173 Phosphorylation Q00535 (CDK5) , P53778 (MAPK12) , P17612 (PRKACA) , P49841 (GSK3B) HPRD   PhosphoSitePlus 19651622, 17287340, 9771888, ...9199504, 12387894, 9565682, 9832145, 9353289, 9735171, 7566346, 20068231
S177 Phosphorylation Q00535 (CDK5) HPRD   PhosphoSitePlus 19651622, 17287340, 9771888, ...7566346, 12387894, 9565682, 9832145, 20068231
S179 Phosphorylation Q16816 (PHKG1) HPRD 19651622, 8999860, 20068231
S180 Phosphorylation HPRD 19651622, 12387894, 20068231
S183 Phosphorylation HPRD 20068231
S200 Phosphorylation Q16512 (PKN1) HPRD 11104762
S204 Phosphorylation Q05655 (PRKCD) , Q16816 (PHKG1) , P49841 (GSK3B) , Q9P0L2 (MARK1) , P17612 (PRKACA) , Q9UQM7 (CAMK2A) HPRD   PRO 9485060, 8999860, 9771888, ...10090741, 12387894, 9832145, 7706316, 9735171
T205 Phosphorylation HPRD 19413330
S231 Phosphorylation Q16512 (PKN1) , P53778 (MAPK12) HPRD 9199504, 11104762
S235 Phosphorylation Q9P0L2 (MARK1) , P49840 (GSK3A) HPRD 7931292, 7706316
S263 Phosphorylation Q16512 (PKN1) HPRD 11104762
S267 Phosphorylation Q9P0L2 (MARK1) , P17612 (PRKACA) , Q9UQM7 (CAMK2A) , Q05655 (PRKCD) HPRD   PhosphoSitePlus 9771888, 10737616, 12387894, ...9832145, 7706316, 8163474, 12435421
T284 Phosphorylation Q00535 (CDK5) HPRD 12387894
Y305 Phosphorylation P00519 (ABL1) HPRD 16014719
S307 Phosphorylation P49841 (GSK3B) , Q00535 (CDK5) HPRD   PRO   PhosphoSitePlus 7566346, 1181841, 12387894, ...9565682, 9832145, 9485060
S311 Phosphorylation P49841 (GSK3B) HPRD 19413330, 7566346, 9832145, ...12387894
T314 Phosphorylation HPRD 9832145, 20068231
S315 Phosphorylation P49841 (GSK3B) , Q00535 (CDK5) HPRD   PRO   PhosphoSitePlus 19413330, 11181841, 7566346, ...9771888, 12387894, 9565682, 9832145, 9485060, 19415658, 19651622
S320 Phosphorylation P17612 (PRKACA) HPRD   PhosphoSitePlus 12435421, 8163474
S323 Phosphorylation HPRD 9832145, 12387894
S324 Phosphorylation P49841 (GSK3B) HPRD 9565682, 12387894
S327 Phosphorylation HPRD 15345747
S333 Phosphorylation HPRD 9832145, 12387894
Site PTM Type PTM Enzyme Score Source PMID
S37 Phosphorylation PhosphoSitePlus
T41 Phosphorylation PhosphoSitePlus
S104 Phosphorylation PhosphoSitePlus
S105 Phosphorylation PhosphoSitePlus
S108 Phosphorylation PhosphoSitePlus
T137 Phosphorylation PhosphoSitePlus
S141 Phosphorylation PhosphoSitePlus
S199 Phosphorylation PhosphoSitePlus
S271 Phosphorylation PhosphoSitePlus
S279 Phosphorylation PhosphoSitePlus
S284 Phosphorylation PhosphoSitePlus
S288 Phosphorylation PhosphoSitePlus
S297 Phosphorylation PhosphoSitePlus
Site PTM Type PTM Enzyme Score Source PMID
S170 Phosphorylation PRO 9485060
S173 Phosphorylation PRO 9485060
T176 Phosphorylation PRO 9485060
S233 Phosphorylation PRO 9485060
S336 Phosphorylation PRO 9485060
S344 Phosphorylation PRO 9485060
Site PTM Type PTM Enzyme Score Source PMID
S113 Phosphorylation PhosphoSitePlus
S131 Phosphorylation PhosphoSitePlus
T135 Phosphorylation PhosphoSitePlus
T181 Phosphorylation PhosphoSitePlus
S184 Phosphorylation PhosphoSitePlus
S195 Phosphorylation neXtProt   PhosphoSitePlus 9771888
S198 Phosphorylation neXtProt   PhosphoSitePlus 9771888
S199 Phosphorylation neXtProt   PRO   PhosphoSitePlus 9485060, 9771888
S202 Phosphorylation neXtProt   PRO   PhosphoSitePlus 9485060, 9771888
T205 Phosphorylation neXtProt   PRO   PhosphoSitePlus 9485060, 9771888
T212 Phosphorylation neXtProt 17078951
S214 Phosphorylation O75582 (RPS6KA5) neXtProt 17512525
T231 Phosphorylation neXtProt   PhosphoSitePlus 17078951, 9771888
S235 Phosphorylation neXtProt   PhosphoSitePlus 15994305, 9771888
S262 Phosphorylation O75582 (RPS6KA5) neXtProt   PRO   PhosphoSitePlus 9485060, 17512525, 9771888
S325 Phosphorylation neXtProt   PhosphoSitePlus 9771888
S365 Phosphorylation PRO   PhosphoSitePlus 9485060
S369 Phosphorylation PhosphoSitePlus
S373 Phosphorylation neXtProt   PRO   PhosphoSitePlus 9485060, 9771888
Site PTM Type PTM Enzyme Score Source PMID
S55 Phosphorylation PhosphoSitePlus
T65 Phosphorylation HPRD 12387894
S73 Phosphorylation HPRD   PhosphoSitePlus 17287340
T77 Phosphorylation HPRD   PhosphoSitePlus 17287340
T95 Phosphorylation P06493 (CDK1) HPRD 9614189
T117 Phosphorylation P28482 (MAPK1) HPRD 9832145, 10737616
T123 Phosphorylation P53778 (MAPK12) , P49841 (GSK3B) , P28482 (MAPK1) HPRD   PhosphoSitePlus 11181841, 7566346, 9771888, ...9832145, 18510355, 9199504
S126 Phosphorylation P49841 (GSK3B) HPRD 12387894, 9771888
S137 Phosphorylation P17612 (PRKACA) , P49841 (GSK3B) HPRD 11181841, 12387894, 9771888
S140 Phosphorylation P49841 (GSK3B) , P17612 (PRKACA) HPRD   PRO 9832145, 7566346, 9485060, ...12387894, 9771888
S141 Phosphorylation P17612 (PRKACA) , P49841 (GSK3B) HPRD 19413330, 9565682, 7566346, ...12387894, 9771888
S144 Phosphorylation Q00535 (CDK5) , P49841 (GSK3B) , P28482 (MAPK1) , P53778 (MAPK12) HPRD   PRO   PhosphoSitePlus 19413330, 9771888, 7566346, ...12226093, 1330687, 12387894, 19415658, 9565682, 9832145, 9485060, 9199504
T147 Phosphorylation Q00535 (CDK5) , P49841 (GSK3B) , Q16512 (PKN1) , P53778 (MAPK12) , P17612 (PRKACA) HPRD   PRO   PhosphoSitePlus 19413330, 9546672, 12226093, ...19562805, 9199504, 11181841, 12387894, 9565682, 9832145, 9485060, 9771888, 12435421
S150 Phosphorylation P49841 (GSK3B) HPRD 19562805, 11311121
S152 Phosphorylation HPRD 9832145
T154 Phosphorylation P17612 (PRKACA) , Q00535 (CDK5) , P51812 (RPS6KA3) , P49841 (GSK3B) HPRD 9546672, 11181841, 12387894, ...11311121, 9832145, 9353289
S156 Phosphorylation Q00535 (CDK5) HPRD 9614189
T159 Phosphorylation P49841 (GSK3B) , Q00535 (CDK5) HPRD 7566346, 9832145, 12387894, ...11181841
T173 Phosphorylation P53778 (MAPK12) , P49841 (GSK3B) , Q00535 (CDK5) , P17612 (PRKACA) HPRD   PhosphoSitePlus 19651622, 17287340, 9771888, ...9199504, 12387894, 9565682, 9832145, 9353289, 9735171, 7566346, 20068231
S177 Phosphorylation Q00535 (CDK5) HPRD   PhosphoSitePlus 19651622, 17287340, 9771888, ...7566346, 12387894, 9565682, 9832145, 20068231
S179 Phosphorylation Q16816 (PHKG1) HPRD 19651622, 8999860, 20068231
S180 Phosphorylation HPRD 19651622, 12387894, 20068231
S183 Phosphorylation HPRD 20068231
S200 Phosphorylation Q16512 (PKN1) HPRD 11104762
S204 Phosphorylation Q05655 (PRKCD) , Q9P0L2 (MARK1) , Q9UQM7 (CAMK2A) , P49841 (GSK3B) , Q16816 (PHKG1) , P17612 (PRKACA) HPRD   PRO   PhosphoSitePlus 8999860, 9771888, 10090741, ...12387894, 9832145, 9485060, 7706316, 9735171
T205 Phosphorylation HPRD 19413330
S235 Phosphorylation Q9P0L2 (MARK1) HPRD 7706316
S247 Phosphorylation Q9P0L2 (MARK1) HPRD 7706316
S262 Phosphorylation Q16512 (PKN1) , P53778 (MAPK12) HPRD 9199504, 11104762
S266 Phosphorylation Q9P0L2 (MARK1) , P49840 (GSK3A) HPRD 7931292, 7706316
S294 Phosphorylation Q16512 (PKN1) HPRD 11104762
S298 Phosphorylation Q9P0L2 (MARK1) , P17612 (PRKACA) , Q05655 (PRKCD) , Q9UQM7 (CAMK2A) HPRD   PhosphoSitePlus 9771888, 10737616, 12387894, ...9832145, 7706316, 12435421
T315 Phosphorylation Q00535 (CDK5) HPRD 12387894
Y336 Phosphorylation P00519 (ABL1) HPRD 16014719
S338 Phosphorylation Q00535 (CDK5) , P49841 (GSK3B) HPRD   PRO   PhosphoSitePlus 7566346, 1181841, 12387894, ...9565682, 9832145, 9485060
S342 Phosphorylation P49841 (GSK3B) HPRD 19413330, 7566346, 9832145, ...12387894
T345 Phosphorylation HPRD 9832145, 20068231
S346 Phosphorylation Q00535 (CDK5) , P49841 (GSK3B) HPRD   PRO   PhosphoSitePlus 19413330, 11181841, 7566346, ...9771888, 12387894, 9565682, 9832145, 9485060, 19415658, 19651622
S351 Phosphorylation P17612 (PRKACA) HPRD 12435421
S354 Phosphorylation HPRD 9832145, 12387894
S355 Phosphorylation HPRD 9565682, 12387894
S358 Phosphorylation HPRD 15345747
S364 Phosphorylation HPRD 9832145, 12387894
Site PTM Type PTM Enzyme Score Source PMID
S68 Phosphorylation PhosphoSitePlus
T82 Phosphorylation PhosphoSitePlus
S84 Phosphorylation PhosphoSitePlus
S102 Phosphorylation PhosphoSitePlus
T106 Phosphorylation PhosphoSitePlus
T152 Phosphorylation P49840 (GSK3A) neXtProt 17229088
S170 Phosphorylation PRO 9485060
S173 Phosphorylation P49840 (GSK3A) neXtProt   PRO 17229088, 9485060
T176 Phosphorylation P49840 (GSK3A) neXtProt   PRO 17229088, 9485060
T183 Phosphorylation IntAct 17512525
S185 Phosphorylation IntAct 17512525
Q215 Phosphorylation IntAct 17512525
S233 Phosphorylation PRO 9485060
S367 Phosphorylation PRO 9485060
S375 Phosphorylation PRO 9485060
S404 Phosphorylation neXtProt 17078951
Site PTM Type PTM Enzyme Score Source PMID
T17 Phosphorylation PhosphoSitePlus
Y18 Phosphorylation P06241 (FYN) , P12931 (SRC) PhosphoSitePlus 16115884, 14999081, 15356202
Y29 Phosphorylation PhosphoSitePlus
T30 Phosphorylation PhosphoSitePlus
T39 Phosphorylation PhosphoSitePlus
K44 Methylation PhosphoSitePlus
S46 Phosphorylation P28482 (MAPK1) , P49841 (GSK3B) , P27361 (MAPK3) HPRD   PhosphoSitePlus 18510355, 9614189
T50 Phosphorylation P27361 (MAPK3) , P28482 (MAPK1) , P49841 (GSK3B) , Q15759 (MAPK11) , O15264 (MAPK13) , P53778 (MAPK12) , Q16539 (MAPK14) HPRD   neXtProt   PhosphoSitePlus 18510355, 15632108, 9614189
T52 Phosphorylation PhosphoSitePlus
S56 Phosphorylation PhosphoSitePlus
S68 Phosphorylation PhosphoSitePlus
T69 Phosphorylation O15264 (MAPK13) , P53778 (MAPK12) , Q16539 (MAPK14) , Q15759 (MAPK11) neXtProt   PhosphoSitePlus 15632108
T71 Phosphorylation PhosphoSitePlus
T95 Phosphorylation PhosphoSitePlus
T101 Phosphorylation PhosphoSitePlus
T102 Phosphorylation PhosphoSitePlus
T111 Phosphorylation PhosphoSitePlus
S113 Phosphorylation PhosphoSitePlus
T123 O-Glycosylation PhosphoSitePlus
T123 Phosphorylation HPRD   PhosphoSitePlus 12387894
K130 Methylation PhosphoSitePlus
S131 Phosphorylation HPRD   PhosphoSitePlus 17287340
K132 Methylation PhosphoSitePlus
T135 Phosphorylation HPRD   PhosphoSitePlus 17287340
T149 Phosphorylation Q5S007 (LRRK2) PhosphoSitePlus 24113872
T153 Phosphorylation P06493 (CDK1) , Q16539 (MAPK14) , P28482 (MAPK1) , O15264 (MAPK13) , Q5S007 (LRRK2) , P27361 (MAPK3) , P49841 (GSK3B) , Q00535 (CDK5) , Q15759 (MAPK11) , P53778 (MAPK12) HPRD   neXtProt   PhosphoSitePlus 10090741, 24113872, 15632108, ...9614189
K163 Acetylation PhosphoSitePlus
K163 Methylation PhosphoSitePlus
T169 Phosphorylation PhosphoSitePlus
K174 Acetylation PhosphoSitePlus
K174 Methylation PhosphoSitePlus
K174 Ubiquitination PhosphoSitePlus
T175 Phosphorylation P28482 (MAPK1) , P49841 (GSK3B) , P27361 (MAPK3) HPRD   PhosphoSitePlus 10090741, 9832145, 9614189, ...10737616
K180 Methylation PhosphoSitePlus
T181 Phosphorylation P49841 (GSK3B) , P45984 (MAPK9) , P45983 (MAPK8) , P53778 (MAPK12) , P27361 (MAPK3) , Q00535 (CDK5) , P28482 (MAPK1) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 12801934, 11181841, ...7566346, 9771888, 9614189, 12387894, 18510355, 15994305, 9832145, 17078951, 9199504
S184 Phosphorylation PhosphoSitePlus
S185 Phosphorylation PhosphoSitePlus
S191 Phosphorylation PhosphoSitePlus
S195 Phosphorylation P49841 (GSK3B) , P17612 (PRKACA) HPRD   PhosphoSitePlus 11181841, 20679343, 12387894, ...9771888
Y197 Phosphorylation Q5TCY1 (TTBK1) , P42684 (ABL2) PRO   PhosphoSitePlus 16923168, 20110615
S198 Phosphorylation Q5TCY1 (TTBK1) , P17612 (PRKACA) , P49841 (GSK3B) HPRD   PRO   PhosphoSitePlus 9771888, 16923168, 12387894, ...9832145, 7566346
S199 Phosphorylation P27361 (MAPK3) , P28482 (MAPK1) , Q13627 (DYRK1A) , P49841 (GSK3B) , Q00535 (CDK5) , Q5TCY1 (TTBK1) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 25918155, 20679343, ...9771888, 16923168, 19413330, 9614189, 12387894, 16257959, 9565682, 9485060, 17078951, 20308788, 7566346, 10090668
S202 Phosphorylation P45984 (MAPK9) , Q13627 (DYRK1A) , Q00535 (CDK5) , P48730 (CSNK1D) , Q15759 (MAPK11) , Q5TCY1 (TTBK1) , P28482 (MAPK1) , P06493 (CDK1) , P45983 (MAPK8) , P53778 (MAPK12) , Q16539 (MAPK14) , P27361 (MAPK3) , P49841 (GSK3B) , O15264 (MAPK13) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 9771888, 10090741, ...17906291, 7566346, 1330687, 12387894, 9485060, 15632108, 10090668, 9832145, 9199504, 19413330, 14761950, 16923168, 9614189, 15994305, 26555036, 12801934, 12226093, 9565682, 17078951, 20308788, 19415658
T205 Phosphorylation P06493 (CDK1) , P49841 (GSK3B) , Q15759 (MAPK11) , P53778 (MAPK12) , Q5S007 (LRRK2) , O15264 (MAPK13) , Q00535 (CDK5) , P48730 (CSNK1D) , P28482 (MAPK1) , P27361 (MAPK3) , Q13627 (DYRK1A) , Q16539 (MAPK14) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 24113872, 9771888, ...19562805, 12387894, 9485060, 15632108, 10090668, 20679343, 11181841, 10090741, 19413330, 14761950, 9199504, 9614189, 15994305, 26555036, 25918155, 12226093, 9565682, 17078951, 20308788
S208 O-Glycosylation PhosphoSitePlus
S208 Phosphorylation P49841 (GSK3B) HPRD   PhosphoSitePlus 19562805, 11311121
S210 Phosphorylation P49841 (GSK3B) HPRD   PhosphoSitePlus 9832145, 20679343
T212 Phosphorylation O15264 (MAPK13) , Q92630 (DYRK2) , Q13627 (DYRK1A) , Q9P0L2 (MARK1) , P27361 (MAPK3) , P28482 (MAPK1) , P49841 (GSK3B) , P17612 (PRKACA) , P45984 (MAPK9) , P51812 (RPS6KA3) , P06493 (CDK1) , Q00535 (CDK5) , P23443 (RPS6KB1) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 25918155, 9546672, ...16987243, 11181841, 10090741, 17906291, 16364302, 9614189, 12387894, 11311121, 15542598, 9832145, 9353289, 17078951, 21985311, 20308788, 10090668
S214 Phosphorylation Q00535 (CDK5) , Q13131 (PRKAA1) , P17612 (PRKACA) , P23443 (RPS6KB1) , Q9P0L2 (MARK1) , O00141 (SGK1) , P49841 (GSK3B) , Q16512 (PKN1) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 16982696, 20679343, ...11104762, 9546672, 16364302, 10090741, 9614189, 12387894, 9832145, 21204788, 17078951, 16443603, 12435421, 16987243, 10090668, 21985311
T217 Phosphorylation Q00535 (CDK5) , Q9P0L2 (MARK1) , O15264 (MAPK13) , P27361 (MAPK3) , P28482 (MAPK1) , P45984 (MAPK9) , Q13627 (DYRK1A) , P49841 (GSK3B) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 25918155, 16987243, ...11181841, 10090741, 7566346, 9771888, 9614189, 12387894, 9832145, 17078951, 16443603, 21985311, 20308788, 9199504
T220 Phosphorylation PhosphoSitePlus
T231 Phosphorylation P27361 (MAPK3) , P53778 (MAPK12) , P06493 (CDK1) , P28482 (MAPK1) , P49840 (GSK3A) , Q9P0L2 (MARK1) , P45984 (MAPK9) , P49841 (GSK3B) , P45983 (MAPK8) , Q00535 (CDK5) , Q13131 (PRKAA1) HPRD   neXtProt   PRO   PhosphoSitePlus 18052981, 9771888, 10090741, ...7566346, 12387894, 9735171, 21204788, 9832145, 16443603, 18055457, 9199504, 9614189, 17045592, 19651622, 9353289, 17287340, 12801934, 9565682, 15542598, 25573097, 17078951, 20068231
S235 Phosphorylation O15264 (MAPK13) , P28482 (MAPK1) , P27361 (MAPK3) , P53778 (MAPK12) , P06493 (CDK1) , Q00535 (CDK5) , Q16539 (MAPK14) , P49841 (GSK3B) , Q15759 (MAPK11) , P49840 (GSK3A) HPRD   neXtProt   PRO   PhosphoSitePlus 19651622, 17287340, 9771888, ...10090741, 7566346, 9614189, 12387894, 17045592, 9565682, 15994305, 9832145, 20068231, 16443603, 9735171, 15632108, 26555036
S237 Phosphorylation Q16816 (PHKG1) HPRD   phospho.ELM   PhosphoSitePlus 19651622, 8999860, 20068231
S238 O-Glycosylation PhosphoSitePlus
S238 Phosphorylation HPRD   PRO   PhosphoSitePlus 19651622, 16443603, 12387894, ...20068231
S241 Phosphorylation HPRD   PhosphoSitePlus 20068231
T245 Phosphorylation P17612 (PRKACA) PhosphoSitePlus 10090741
K254 Methylation PhosphoSitePlus
K254 Ubiquitination PhosphoSitePlus
S258 Phosphorylation P41743 (PRKCI) , Q16512 (PKN1) , Q05513 (PRKCZ) , P17252 (PRKCA) , Q02156 (PRKCE) , P05771-2 (PRKCB) , P05129 (PRKCG) HPRD   PhosphoSitePlus 11104762
K259 Acetylation PhosphoSitePlus
S262 Phosphorylation O14757 (CHEK1) , P23443 (RPS6KB1) , Q16816 (PHKG1) , Q05655 (PRKCD) , Q9UQM7 (CAMK2A) , P17612 (PRKACA) , Q7KZI7 (MARK2) , P49841 (GSK3B) , P19525 (EIF2AK2) , O96017 (CHEK2) , Q9P0L2 (MARK1) , P49840 (GSK3A) , Q00535 (CDK5) , Q13131 (PRKAA1) HPRD   IntAct   neXtProt   phospho.ELM   PRO   PhosphoSitePlus 18052981, 9771888, 10090741, ...12387894, 23001711, 20067632, 17360912, 12435421, 9735171, 17573348, 18424437, 20159774, 20308788, 21204788, 9832145, 7706316, 16443603, 8999860, 16364302, 9614189, 17045592, 21985244, 9485060, 15652175, 17078951, 21311567
T263 Phosphorylation HPRD   PhosphoSitePlus 19413330
K267 Methylation PhosphoSitePlus
K274 Acetylation PhosphoSitePlus
K280 Acetylation PhosphoSitePlus
K281 Acetylation PhosphoSitePlus
S285 Phosphorylation phospho.ELM   PhosphoSitePlus 8999860
S289 Phosphorylation PhosphoSitePlus
K290 Acetylation PhosphoSitePlus
K290 Methylation PhosphoSitePlus
S293 Phosphorylation Q02156 (PRKCE) , P05771 (PRKCB) , P17612 (PRKACA) , P17252 (PRKCA) , P41743 (PRKCI) , Q05513 (PRKCZ) , P05129 (PRKCG) , Q9P0L2 (MARK1) HPRD   PhosphoSitePlus 7706316, 11104762, 9614189
K298 Acetylation PhosphoSitePlus
S305 Phosphorylation P17252 (PRKCA) , Q9P0L2 (MARK1) , P17612 (PRKACA) HPRD   phospho.ELM   PhosphoSitePlus 8999860, 7706316
Y310 Phosphorylation P42684 (ABL2) PhosphoSitePlus 20110615
K311 Acetylation PhosphoSitePlus
K311 Ubiquitination PhosphoSitePlus
S316 Phosphorylation P17612 (PRKACA) PhosphoSitePlus 10090741
K317 Methylation PhosphoSitePlus
S320 Phosphorylation P53778 (MAPK12) , Q9P0L2 (MARK1) , P17612 (PRKACA) , Q16512 (PKN1) HPRD   PhosphoSitePlus 10090741, 9199504, 11104762
K321 Acetylation PhosphoSitePlus
S324 Phosphorylation P17612 (PRKACA) , P05771 (PRKCB) , Q9P0L2 (MARK1) , P05129 (PRKCG) , Q05513 (PRKCZ) , P49840 (GSK3A) , P17252 (PRKCA) , P41743 (PRKCI) , Q02156 (PRKCE) HPRD   PhosphoSitePlus 11104762, 10090741, 9614189, ...7706316, 7931292
K331 Acetylation PhosphoSitePlus
K340 Sumoylation PhosphoSitePlus
S341 Phosphorylation PhosphoSitePlus
K343 Acetylation PhosphoSitePlus
K343 Ubiquitination PhosphoSitePlus
R349 Methylation PhosphoSitePlus
S352 Phosphorylation P05771 (PRKCB) , Q16512 (PKN1) , P17252 (PRKCA) , Q02156 (PRKCE) , P05129 (PRKCG) , Q05513 (PRKCZ) , P41743 (PRKCI) HPRD   phospho.ELM   PhosphoSitePlus 8999860, 11104762
K353 Acetylation PhosphoSitePlus
K353 Ubiquitination PhosphoSitePlus
S356 O-Glycosylation PhosphoSitePlus
S356 Phosphorylation Q05655 (PRKCD) , Q9P0L2 (MARK1) , P17612 (PRKACA) , Q7KZI7 (MARK2) , Q9UQM7 (CAMK2A) , P19525 (EIF2AK2) , Q13131 (PRKAA1) HPRD   neXtProt   PRO   PhosphoSitePlus 9771888, 10090741, 10737616, ...9614189, 12387894, 9832145, 20067632, 7706316, 21204788, 20658989, 12435421
T361 Phosphorylation PhosphoSitePlus
K369 Acetylation PhosphoSitePlus
K369 Ubiquitination PhosphoSitePlus
T373 Phosphorylation Q00535 (CDK5) HPRD   PhosphoSitePlus 12387894
K375 Ubiquitination PhosphoSitePlus
T377 Phosphorylation PhosphoSitePlus
T386 Phosphorylation PhosphoSitePlus
Y394 Phosphorylation P42684 (ABL2) , P00519 (ABL1) HPRD   PhosphoSitePlus 20110615, 16014719
S396 Phosphorylation P27361 (MAPK3) , P53778 (MAPK12) , Q13627 (DYRK1A) , P45983 (MAPK8) , P28482 (MAPK1) , O15264 (MAPK13) , Q9P0L2 (MARK1) , P48730 (CSNK1D) , Q13131 (PRKAA1) , Q00535 (CDK5) , P49841 (GSK3B) , P49840 (GSK3A) HPRD   IntAct   neXtProt   PRO   PhosphoSitePlus 18052981, 10090741, 7566346, ...10818091, 12387894, 9485060, 20679343, 21204788, 9832145, 16443603, 21985311, 18055457, 14761950, 9614189, 16478768, 10620503, 25918155, 21985244, 1181841, 9565682, 15542598, 17078951, 20080974, 20308788
S400 O-Glycosylation PhosphoSitePlus
S400 Phosphorylation P49841 (GSK3B) HPRD   PhosphoSitePlus 19413330, 20679343, 7566346, ...12387894, 9832145, 16478768
T403 Phosphorylation HPRD   PhosphoSitePlus 9832145, 20068231
S404 Phosphorylation Q16539 (MAPK14) , O15264 (MAPK13) , P53778 (MAPK12) , P48730 (CSNK1D) , P28482 (MAPK1) , P06493 (CDK1) , Q15759 (MAPK11) , P49841 (GSK3B) , P49840 (GSK3A) , Q13627 (DYRK1A) , Q00535 (CDK5) , P27361 (MAPK3) HPRD   IntAct   neXtProt   PRO   PhosphoSitePlus 18052981, 9771888, 10090741, ...17906291, 7566346, 10818091, 12387894, 9485060, 15632108, 20679343, 11181841, 16257959, 9832145, 18055457, 19413330, 14761950, 9614189, 19651622, 15994305, 10620503, 25918155, 21985244, 9565682, 15542598, 17078951, 20080974, 20308788, 19415658
S409 Phosphorylation P17612 (PRKACA) , P49841 (GSK3B) , Q00535 (CDK5) HPRD   neXtProt   PhosphoSitePlus 10090741, 12435421, 17078951, ...18052981
S412 Phosphorylation HPRD   PhosphoSitePlus 9832145, 12387894
S413 Phosphorylation P49841 (GSK3B) HPRD   PhosphoSitePlus 9565682, 12387894
T414 Phosphorylation PhosphoSitePlus
S416 Phosphorylation Q5TCY1 (TTBK1) , P17612 (PRKACA) , Q9UQM7 (CAMK2A) HPRD   PRO   PhosphoSitePlus 10090741, 16923168, 16000144, ...15345747
S422 Phosphorylation P49841 (GSK3B) , Q5TCY1 (TTBK1) , P27361 (MAPK3) , Q15759 (MAPK11) , P53778 (MAPK12) , Q9P0L2 (MARK1) , Q16539 (MAPK14) , Q13627 (DYRK1A) , O15264 (MAPK13) , P28482 (MAPK1) , Q00535 (CDK5) , Q13131 (PRKAA1) HPRD   neXtProt   PRO   PhosphoSitePlus 25918155, 10090741, 16923168, ...9614189, 21204788, 9832145, 12387894, 17078951, 15632108
T427 Phosphorylation Q5TCY1 (TTBK1) PRO   PhosphoSitePlus 16923168
S433 Phosphorylation PhosphoSitePlus
S435 Phosphorylation PhosphoSitePlus


Proteoforms

PRO ID (Short Label) Sites PTM Enzyme Source PMID
PR:000027682 (hMAPT/iso:Fetal-tau/Phos:7) pS141,pS144,pT147,pS204,pS307,pS315 PRO 9485060
PR:000027680 (hMAPT/iso:Tau-B/Phos:7) pS170,pS173,pT176,pS233,pS336,pS344 PRO 9485060
PR:000027678 (hMAPT/iso:Tau-C/Phos:7) pS199,pS202,pT205,pS262,pS365,pS373 PRO 9485060
PR:000027676 (hMAPT/iso:Tau-D/Phos:7) pS140,pS144,pT147,pS204,pS338,pS346 PRO 9485060
PR:000027674 (hMAPT/iso:Tau-E/Phos:7) pS170,pS173,pT176,pS233,pS367,pS375 PRO 9485060
PR:000027430 (hMAPT/iso:Tau-F/Phos:3) pY197,pS198 PRO 16923168
PR:000027448 (hMAPT/iso:Tau-F/Phos:1) pT181 PRO
PR:000027450 (hMAPT/iso:Tau-F/Phos:2) pT231 PRO 16443603
PR:000027453 (hMAPT/iso:Tau-F/Phos:4) pS199,pS202 PRO 16923168
PR:000027456 (hMAPT/iso:Tau-F/Phos:5) pS422,pT427 PRO 16923168
PR:000027459 (hMAPT/iso:Tau-F/Phos:6) pS416,pS422,pT427 PRO 16923168
PR:000027631 (hMAPT/Phos:7) pS199,pS202,pT205,pS262,pS396,pS404 PRO 9485060
PR:000027633 (hMAPT/Phos:8) pS262 PRO 16443603
PR:000027635 (hMAPT/Phos:9) pS396 PRO 16443603
PR:000027638 (hMAPT/Phos:10) pT217 PRO 16443603
PR:000027640 (hMAPT/Phos:11) pS214,pT217 PRO 16443603
PR:000027642 (hMAPT/Phos:12) pT231 PRO 16443603
PR:000027645 (hMAPT/Phos:16) pS202,pT205 PRO 19413330
PR:000027647 (hMAPT/Phos:17) pS202 PRO 19413330
PR:000027649 (hMAPT/Phos:18) pS199,pS202,pT205,pT212,pS214 PRO 10090668
PR:000027656 (hMAPT/iso:Tau-F/Phos:7) pS199,pS202,pT205,pS262,pS396,pS404 PRO 9485060
PR:000027658 (hMAPT/iso:Tau-F/Phos:8) pS262 PRO 16443603
PR:000027660 (hMAPT/iso:Tau-F/Phos:9) pS396 PRO 16443603
PR:000027662 (hMAPT/iso:Tau-F/Phos:10) pT217 PRO 16443603
PR:000027664 (hMAPT/iso:Tau-F/Phos:11) pS214,pT217 PRO 16443603
PR:000027667 (hMAPT/iso:Tau-F/Phos:13) pT231,pS235 PRO 16443603
PR:000027668 (hMAPT/iso:Tau-F/Phos:14) pT231,pS238 PRO 16443603
PR:000027670 (hMAPT/iso:Tau-F/Phos:15) pS356 PRO 20658989
PR:000027672 (hMAPT/iso:Tau-F/Phos:18) pS199,pS202,pT205,pT212,pS214 PRO 10090668


PTM sites affected in variants

Site Variant Source PMID Disease [Sample source]
S171 L171 Biomuta DOID:4159 / skin cancer [ cosmic ]
S177 L177 Biomuta DOID:4159 / skin cancer [ cosmic ]
S180 F180 Biomuta DOID:4159 / skin cancer [ cosmic ]
T205 N205 Biomuta DOID:4606 / bile duct cancer [ cosmic ] DOID:10534 / stomach cancer [ cosmic, icgc ]
G205 R205 Biomuta DOID:4159 / skin cancer [ cosmic ]
G205 V205 Biomuta DOID:1324 / lung cancer [ cosmic, icgc ]
G205 E205 Biomuta [ cosmic ]
P212 S212 Biomuta DOID:4159 / skin cancer [ cosmic ]
P212 L212 Biomuta DOID:4159 / skin cancer [ cosmic, icgc ] DOID:8618 / oral cavity cancer [ cosmic ] DOID:11934 / head and neck cancer [ icgc ]
S232 Y232 Biomuta DOID:8618 / oral cavity cancer [ cosmic ] DOID:11934 / head and neck cancer [ icgc ]
S232 F232 Biomuta DOID:4159 / skin cancer [ cosmic ]
S238 P238 Biomuta DOID:3571 / liver cancer [ cosmic ]
S263 L263 Biomuta DOID:363 / uterine cancer [ cosmic, icgc ] DOID:10534 / stomach cancer [ cosmic ]
S267 F267 Biomuta DOID:1324 / lung cancer [ cosmic, icgc ]
S320 N320 Biomuta DOID:363 / uterine cancer [ icgc ]
S396 F396 Biomuta DOID:4159 / skin cancer [ cosmic ]
T466 M466 Biomuta DOID:1324 / lung cancer [ cosmic ]
S508 * Biomuta DOID:263 / kidney cancer [ cosmic ]
S552 L552 Biomuta DOID:4159 / skin cancer [ cosmic ]
S555 F555 Biomuta DOID:4159 / skin cancer [ cosmic ]
K571 E571 Biomuta DOID:9256 / colorectal cancer [ cosmic ]
T580 N580 Biomuta DOID:4606 / bile duct cancer [ cosmic ] DOID:10534 / stomach cancer [ cosmic, icgc ]
S602 G602 Biomuta DOID:1781 / thyroid gland cancer [ cosmic ] DOID:1781 / thyroid cancer [ icgc ]
S610 L610 Biomuta DOID:4159 / skin cancer [ cosmic ]
S633 N633 Biomuta DOID:9256 / colorectal cancer [ cosmic, icgc ]
S633 I633 Biomuta DOID:363 / uterine cancer [ cosmic, icgc ]
S669 L669 Biomuta DOID:363 / uterine cancer [ cosmic, icgc ] DOID:10534 / stomach cancer [ cosmic ]
K670 N670 Biomuta DOID:9256 / colorectal cancer [ cosmic ]
S673 F673 Biomuta DOID:1324 / lung cancer [ cosmic, icgc ]
S726 N726 Biomuta DOID:363 / uterine cancer [ icgc ]
T744 M744 Biomuta DOID:4362 / cervical cancer [ cosmic, icgc ]